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14 results for 'IR.HIR.KFaR.FoR'

   IR.HIR.KFaR.FoR  Foliose red seaweeds on exposed lower infralittoral rock

A dense turf of foliose red seaweeds on exposed or moderately exposed lower infralittoral rock, generally, at or below the lower limit of the kelp. Most of the red seaweeds are common to the kelp zone above, while the faunal component of the biotope is made up of species that are found either in the kelp zone or the animal-dominated upper circalittoral below. Foliose species commonly present include Dilsea carnosa, Hypoglossum hypoglossoides, Schottera nicaeensis, Cryptopleura ramosa and Delesseria sanguinea. The red seaweed species composition varies considerably; at some sites a single species may dominate (particularly Plocamium cartilagineum). Small filamentous red seaweeds can be found here as well. These include species such as Heterosiphonia plumosa, Vertebrata byssoides. As well as a varied red seaweed component, this biotope may also contain occasional kelp plants and patches of the brown foliose seaweed Dictyota dichotoma. Coralline crusts covers the bedrock beneath the seaweeds. The fauna generally comprises low-encrusting forms such as the tubeworms Spirobranchus spp., anthozoans including Alcyonium digitatum, Urticina felina and Caryophyllia smithii) and occasional sponge crusts such as Cliona celata, Amphilectus fucorum, Sycon ciliatum and Dysidea fragilis. More mobile fauna include the gastropod Calliostoma zizyphinum, the echinoderms Echinus esculentus as well as the starfish Asterias rubens and Marthasterias glacialis and lastly, the crab Cancer pagurus. Bryozoan crusts such as Electra pilosa can be found fronds on the foliose red seaweeds while scattered hydroids such as Nemertesia antennina form colonies on shells, cobbles and available rock. At some sites erect bryozoans Crisia spp. and Bugula spp. are present. Ascidians such as Clavelina lepadiformis and Clavelina lepadiformis may also be common. In the north the foliose red seaweed Metacallophyllis laciniata may occur.

   IR.HIR.KFaR.LhypRVt  Laminaria hyperborea and red seaweeds on exposed vertical rock

On exposed coasts with moderately strong to weak tidal currents generally at depths of 0-10m, vertical rock communities dominated by frequent Laminaria hyperborea and its commonly associated red seaweeds Delesseria sanguinea, Cryptopleura ramosa and Plocamium cartilagineum can be found. Within this biotope the jewel anemone Corynactis viridis is frequently found in dense aggregations attached to the vertical rock surface. This biotope contains 5 sub-biotopes, distinguished by their biogeography. On the west coast of Scotland, the Northern Isles and the Isle of Man on extremely exposed coasts a variant of this biotope characterised by frequent Metridium senile and occasional Cylista elegans can be found. Further south on the west coast of Ireland, southern Scotland, Wales, and south west England a second variant characterised by frequent Alcyonium digitatum and occasional Cliona celata can be distinguished. A third variant has been recorded from Northern Ireland characterised by the red seaweeds Lithophyllum and Ptilota gunneri, the sea squirt Dendrodoa grossularia and the bryozoan Membranipora membranacea. South from the Isle of Man, on the Welsh Coast, and on the south west and southern English coasts a fourth variant of this biotope is found, which is characterised by the barnacle Balanus crenatus, which may be more frequent in this sub-biotope, and the rarity of Alcyonium digitatum, a species which is more frequent in other variants. This variant has mainly been recorded in shallow water (0-5m). The final biogeographic variant of this biotope is, as with the previous variant, found on the coasts of Wales and south west England. It can be distinguished from the previous variant by the frequent Diplosoma listerianum and occasional Lissoclinum perforatum, although these species are not always present.

   IR.HIR.KFaR.LhypFa  Laminaria hyperborea forest with a faunal cushion (sponges and polyclinids) and foliose red seaweeds on very exposed upper infralittoral rock

Very exposed and exposed, but wave-surged, upper infralittoral bedrock and massive boulders characterised by a dense forest of the kelp Laminaria hyperborea with a high diversity of seaweeds and invertebrates. The shallowest kelp plants are often short or stunted, while deeper plants are taller with heavily epiphytised stipes with foliose red seaweeds such as Delesseria sanguinea, Cryptopleura ramosa or Plocamium cartilagineum or even the brown seaweed Dictyota dichotoma. Also found on the stipes or on the rock below the canopy are red seaweeds including Phycodrys rubens, Kallymenia reniformis, Metacallophyllis laciniata, Caryophyllia smithii, and Corallina officinalis, while encrusting coralline algae can cover any bare patches of rock. At some sites the red seaweeds can be virtually mono-specific, while at other sites show considerable variation containing a dense mixed turf of a large variety of species. The red seaweed Odonthalia dentata can be present in the north. The faunal and floral under-storey is generally rich in species due, in part, to the relatively low urchin-grazing pressure in such shallow exposed conditions. The faunal composition of this biotope varies markedly between sites, but commonly occurring are the soft coral Alcyonium digitatum and the anthozoans Cylista elegans and Corynactis viridis. Sponges form a prominent part of the community with variable amounts of the sponges Halichondria panicea and Pachymatisma johnstonia and several other species. The crab Cancer pagurus and the starfish Asterias rubens are normally present in small numbers foraging beneath the canopy, while the sea urchins Echinus esculentus and Urticina felina graze on the seaweeds. The hydroid Obelia geniculata, the ascidian Botryllus schlosseri and the bryozoan Membranipora membranacea compete for space on the kelp, whereas the bryozoan Electra pilosa also can be found on foliose red seaweeds.

   IR.HIR.KFaR.LhypPar  Sparse Laminaria hyperborea and dense Paracentrotus lividus on exposed infralittoral limestone

This biotope is known from only one location, the Aran Islands, Co. Galway. Here, a limestone platform between 3 m and 6 m of depth is dominated by a dense population of the urchin Paracentrotus lividus, which heavily graze and burrow into the soft limestone. So intense is the grazing pressure that the rock appears completely bare, except for a coralline algal crust and occasional Laminaria hyperborea and Saccorhiza polyschides. The anthozoans Cylista elegans and Corynactis viridis are also present, though at low abundance. The grazed kelp also extends deeper to 20 to 25 m further offshore. (Only one CB record within this biotope).

   IR.HIR.KFaR.AlaAnCrSp  Alaria esculenta forest with dense anemones and crustose sponges on extremely exposed infralittoral bedrock

This biotope has only been recorded from Rockall, where Alaria esculenta appears to replace Laminaria hyperborea as the dominant kelp forest species on the extremely wave-exposed steep and vertical rock, a zone that extends from 14 m down to 35 m. Beneath the A. esculenta canopy, the rock surface is covered by a dense turf of anthozoans such as Cylista elegans, Phellia gausapata and Corynactis viridis, encrusting sponges and coralline algae. The gastropod Margarites helicinus can be found grazing on the kelp fronds, whereas the crab Cancer pagurus can be found among the kelp stipes. The bryozoan Tubularia indivisa also occur, but it does not form such a dense turf as in more shallow waters, while the ascidian Botryllus leachi is found encrusting the large brown seaweeds. Cryptopleura ramosa is the dominant red seaweed on horizontal surfaces. The kelp Laminaria digitata is reported to occur mixed with A. esculenta on the nearby Helen's reef.

   IR.HIR.KFaR.FoR.Dic  Foliose red seaweeds with dense Dictyota dichotoma and/or Dictyopteris polypodioides on exposed lower infralittoral rock

A dense turf of foliose red seaweeds mixed with a dense turf of the foliose brown seaweeds Dictyota dichotoma and/or Dictyopteris polypodioides on exposed and moderately exposed lower infralittoral rock, generally at or below the lower limit of the kelp zone. In some areas the lower infralittoral is subject to a moderate amount of scour from nearby sand. D. dichotoma is relatively tolerant of such scour and in such areas a zone forms with other sand-tolerant seaweeds. D. membranacea is confined to south-western coasts. Typically brown seaweeds dominate the seabed or are at least in equal abundance to the red seaweeds, some of which may also form dense stands such as Plocamium cartilagineum, Calliblepharis ciliata, Cryptopleura ramosa, Bonnemaisonia asparagoides, Heterosiphonia plumosa, Delesseria sanguinea and Vertebrata byssoides. The urchin Echinus esculentus can be found grazing the rock surface which can be covered in coralline algae. The anthozoans Caryophyllia smithii and Alcyonium digitatum are usually present in this biotope along with the tube-building worm Spirobranchus sp. which is more common in sand-scoured areas. The starfish Asterias rubens and Henricia sp. and sponge crusts including Cliona celata can also be found here. D. dichotoma also occurs in the kelp park, and records should only be assigned to this biotope where kelp such as Laminaria hyperborea is sparse or absent and a relatively high density of D. dichotoma and/or D. membranacea is present.

   IR.HIR.KFaR.Ala  Alaria esculenta on exposed sublittoral fringe bedrock

Exposed sublittoral fringe bedrock with an Alaria esculenta forest and an encrusting fauna of the mussel Mytilus edulis and barnacles such as Semibalanus balanoides. The kelp Laminaria digitata can be part of the canopy. Underneath the canopy are red seaweeds such as Mastocarpus stellatus and Palmaria palmata, while encrusting coralline red algae such as Lithothamnion graciale covers the rock surface. The limpet Patella vulgata can be found grazing the rock surface, while the whelk Nucella lapillus is preying on the limpets, barnacles and mussels. Two variants of this biotope are described. In more wave exposed conditions Laminaria digitata is absent and the rock surface is often characterised by dense patches of mussels (Ala.Myt). In slightly less exposed sites the A. esculenta is mixed with L. digitata (Ala.Ldig).

   IR.HIR.KFaR.LhypR  Laminaria hyperborea with dense foliose red seaweeds on exposed infralittoral rock

Very exposed to exposed infralittoral bedrock or large boulders characterised by the kelp Laminaria hyperborea, beneath which is a dense turf of foliose red seaweeds. Three variations of this biotope are described: the upper infralittoral kelp forest (LhypR.Ft), the kelp park below (LhypR.Pk) and a third type of kelp forest, confined to southern England, that is characterised by a mixture of L. hyperborea and Laminaria ochroleuca (LhypR.Loch). The fauna of these biotopes is markedly less abundant than kelp forests in areas of greater wave surge (LhypFa); sponges, anthozoans and polyclinid ascidians may be present, though never at high abundance. Beneath the under-storey of red seaweeds, the rock surface is generally covered with encrusting coralline algae.

   IR.HIR.KFaR  Kelp with cushion fauna and/or foliose red seaweeds

Rocky habitats in the infralittoral zone subject to exposed to extremely exposed wave action or strong tidal streams. Typically the rock supports a community of kelp Laminaria hyperborea with foliose seaweeds and animals, the latter tending to become more prominent in areas of strongest water movement (LhypFa, LhypR and LhypR.Pk). The depth to which the kelp extends varies according to water clarity, exceptionally (e.g. St Kilda) reaching 45 m. In some areas, there may be a band of dense foliose seaweeds (reds or browns) below the main kelp zone (FoR). The sublitttoral fringe is characterised by dabberlocks Alaria esculenta (Ala biotopes). In very strong wave action the sublittoral fringe A. esculenta zone extends to 5 to 10 m depth, whilst at Rockall A. esculenta replaces L. hyperborea as the dominant kelp in the infralittoral zone (AlaAnCrSp).

   IR.HIR.KFaR.Ala.Myt  Alaria esculenta, Mytilus edulis and coralline crusts on very exposed sublittoral fringe bedrock

Very exposed sublittoral fringe bedrock characterised by the kelp Alaria esculenta and dense patches of small individuals of the mussel Mytilus edulis, both of which grow over a dense cover of encrusting coralline algae. Foliose red seaweeds may also be present, but the species composition and their abundance vary between sites. Species such as Corallina officinalis occur widely. The kelp Laminaria digitata is usually absent, although stunted plants may be present at a few sites. The limpet Patella vulgata and the barnacle Semibalanus balanoides are often common. Patches of anthozoans and the hydroid Tubularia spp. occur in more wave-surged areas. In extremely exposed areas the A. esculenta zone can extend as deep as 15 m, where it has less S. balanoides, M. edulis and greater densities of Tubularia spp. (e.g. Barra and shallow areas of Rockall).

   IR.HIR.KFaR.Ala.Ldig  Alaria esculenta and Laminaria digitata on exposed sublittoral fringe bedrock

Exposed sublittoral fringe bedrock characterised by a mixture of the kelps Laminaria digitata and Alaria esculenta with an understorey of red seaweeds including Palmaria palmata and Corallina officinalis with encrusting coralline algal on the rock surface. Anthozoans such as Halichondria panicea, the mussel Mytilus edulis and the barnacle Semibalanus balanoides can be found attached in cracks and crevices. The limpets Patella vulgata or on southern shores Patella ulyssiponensis can be found in their characteristic "scars" grazing the biofilm/algal crusts on the rock surface, while the limpet Patella pellucida is restricted to grazing the kelp fronds. Colonies of the bryozoan Electra pilosa can cover the red seaweeds Mastocarpus stellatus and Chondrus crispus or the rock surface.

   IR.HIR.KFaR.LhypR.Loch  Mixed Laminaria hyperborea and Laminaria ochroleuca forest on exposed infralittoral rock

Mixed Laminaria hyperborea and Laminaria ochroleuca forests on upper infralittoral exposed rock with a dense community of foliose red seaweeds such as Cryptopleura ramosa, and Plocamium cartilagineum as well as small filamentous red seaweeds including Bonnemaisonia asparagoides, Heterosiphonia plumosa, Symphyocladiella parasitica and Vertebrata byssoides. L. hyperborea has a rough stipe which allow dense assemblages of epiphytic red seaweeds to form including the foliose Metacallophyllis laciniata, Delesseria sanguinea and Hypoglossum hypoglossoides. Unlike L. hyperborea, however, L. ochroleuca has a smooth stipe and so it lacks dense assemblages of epiphytic seaweeds L. ochroleuca has a smooth stipe. Encrusting coralline algae often cover much of the rock surface along with a few brown seaweeds including Dictyota dichotoma, Dictyopteris polypodioides and Desmarestia aculeata present as well. In mixed kelp forest L. ochroleuca may predominate with L. hyperborea more common at shallower depths. Whilst foliose red seaweeds dominate the upward-facing rock beneath the kelp canopy, much of the fauna is restricted to crevices or vertical faces, possibly due to grazing pressure. Echinoderms are often common in this biotope, in particular the sea urchin Echinus esculentus and the starfish Asterias rubens and Marthasterias glacialis. Verticals are colonised by anthozoans including the anthozoans Corynactis viridis, Caryophyllia smithii, Actinothoe sphyrodeta and Alcyonium digitatum, while the bryozoan Membranipora membranacea colonise the Laminaria sp. fronds. This biotope is restricted to the coast of Cornwall and the Isles of Scilly. L. ochroleuca occurs at low abundance in other kelp biotopes (sheltered through to exposed) from Dorset to Lundy Island. In such cases, records should be treated as regional variations of these biotopes. Records should only be assigned to the LhypR.Loch biotope when the canopy is dominated by L. ochroleuca alone, or (more usually) by a mixture of both L. hyperborea and L. ochroleuca (at similar abundance). Both this biotope and Lhyp.Loch are common on the Brittany and Normandy coasts.

   IR.HIR.KFaR.LhypR.Pk  Laminaria hyperborea park with dense foliose red seaweeds on exposed lower infralittoral rock

Very exposed to exposed lower infralittoral bedrock or large boulders characterised by a kelp park of Laminaria hyperborea with a dense turf of foliose red seaweeds and encrusting coralline algae. These red seaweeds dominate kelp stipes and bedrock in a similar abundance and composition to the upper infralittoral kelp forest, the most commonly occurring species being Metacallophyllis laciniata, Cryptopleura ramosa, Plocamium cartilagineum, Kallymenia reniformis, Delesseria sanguinea, Phycodrys rubens, Hypoglossum hypoglossoides, Heterosiphonia plumosa and Bonnemaisonia asparagoides. In addition, moderate to high abundance of foliose brown seaweeds, such as Dictyota dichotoma are more common than in the kelp forest above. More upper circalittoral fauna occur in the park than in the kelp forest, such as the cup-coral Caryophyllia smithii. Some species more often present in the kelp park than the forest include the anthozoan Alcyonium digitatum and the featherstar Antedon bifida. The urchin Echinus esculentus, the gastropods Steromphala cineraria and Calliostoma zizyphinum and the starfish Asterias rubens are normally present underneath the canopy along with the anthozoans Urticina felina and Corynactis viridis. The sponge Cliona celata is also present often found boring into shells or soft rock where available. The bryozoan Membranipora membranacea can be found on the L. hyperborea fronds along with the hydroid Obelia geniculata and the ascidian Botryllus schlosseri. The polychaete Spirobranchus sp. is present on the rock surface.

   IR.HIR.KFaR.LhypR.Ft  Laminaria hyperborea forest with dense foliose red seaweeds on exposed upper infralittoral rock

Very exposed to exposed upper infralittoral bedrock or large boulders characterised by a dense forest of the kelp Laminaria hyperborea. On the rock surface beneath the kelp canopy is a dense turf of red foliose seaweeds including Cryptopleura ramosa, Plocamium cartilagineum, Phycodrys rubens and Metacallophyllis laciniata as well as encrusting coralline algae and the foliose brown seaweed Dictyota dichotoma. The red algal turf can be virtually mono-specific, dominated by stands of P. cartilagineum, C. ramosa or Heterosiphonia plumosa, Kallymenia reniformis or in the north, Odonthalia dentata. Other sites may contain a dense mixed turf of these and other species. The dense turf is due, in part, to the relatively low grazing pressure from the urchin Echinus esculentus in such shallow exposed conditions. The shallowest kelp plants are often short or stunted, while deeper plants are taller and the stipes are heavily epiphytised by red seaweeds such as Delesseria sanguinea and Membranoptera alata. The bryozoan Electra pilosa can form colonies on the foliose red seaweeds, while the bryozoan Membranipora membranacea more often can be found on the L. hyperborea fronds along with the ascidian Botryllus schlosseri and the hydroid Obelia geniculata. The gastropods Steromphala cineraria and Calliostoma zizyphinum are found grazing among the kelp holdfasts, while a few individuals of the barnacle Balanus crenatus can present along with the white calcareous tubes of the polychaete Spirobranchus triqueter, where substratum is available. The starfish Asterias rubens can be found predating on polychaetes, mussels and small crustaceans. The soft coral Alcyonium digitatum can be present covering the rock surface as well as the anthozoan Urticina felina.
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