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Modiolus beds on mixed substrata (cobbles, pebbles and coarse muddy sediments) in moderately strong currents or wave exposed areas, typically on the open coast but also in tide-swept channels of marine inlets. Ophiothrix fragilis are often common in this biotope along with the calcareous tubes of Spirobranchus triqueter, anemones such as Alcyonium digitatum and Urticina felina and hydroids such as Abietinaria abietina and Sertularia argentea. Buccinum undatum may also be important and in some areas the clam Mimachlamys varia may be frequent but not in the same abundances as in SS.SBR.SMus.ModMvar. Little information on the infaunal component is given here although it is likely that it is very rich and may highlight more subtle differences in the Modiolus biotopes. This biotope is typified by examples off the north-west Lleyn Peninsula in north Wales and off Co. Down, Northern Ireland.

Muddy gravels and coarse sands in deeper water of continental seas may contain venerid bivalves with beds of Modiolus modiolus. The clumping of the byssus threads of the M. modiolus creates a stable habitat that attracts a very rich infaunal community with a high density of polychaete species including Glycera lapidum, Paradoneis lyra, Aonides paucibranchiata, Laonice bahusiensis, Protomystides bidentata, Lumbrineris spp., Mediomastus fragilis and syllids such as Exogone spp. and Sphaerosyllis spp. Bivalves such as Spisula elliptica, Timoclea ovata and other venerid species are also common. Brittlestars such as Amphipholis squamata may also occur with this community. This biotope is very similar to SS.SMx.OMx.PoVen and the 'boreal off-shore gravel association' and the 'deep Venus community' described by previous workers (Ford 1923; Jones 1951). Similar Modiolus beds (though with a less diverse infauna) on open coast stable boulders, cobbles and sediment are described under SS.SBR.SMus.ModT.