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In sheltered muddy mixed sediments in estuaries or marine inlets with variable or reduced/low salinity communities characterised by Aphelochaeta marioni and Polydora ciliata may be present. Other important taxa may include the polychaetes Nephtys hombergii, Chaetozone zetlandica and Melinna palmata, tubificid oligochaetes and bivalves such as Abra nitida. Conspicuous epifauna may include members of the bivalve family Cardiidae (cockles) and the slipper limpet Crepidula fornicata. This biotope is often found in polyhaline waters.