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Mainly on the mid and lower shore on wave-exposed or moderately wave-exposed coasts of medium and fine sand, sometimes with a fraction of coarse sand, which remains damp throughout the tidal cycle and contains little organic matter. The sediment is often rippled and typically lacks an anoxic sub-surface layer. The infauna is dominated by burrowing amphipods, most notably Pontocrates arenarius, as well as Bathyporeia pelagica, Haustorius arenarius and the isopod Eurydice pulchra. The polychaete fauna is poor, dominated by Scolelepis (Scolelepis) squamata, which tolerates the exposed and mobile sediment conditions. The presence of polychaetes may be seen as coloured burrows running down from the surface of the sediment.